全球人体发现的寄生虫虫种及其生物学分类张进顺1,张颖骞21.河北北方学院寄生虫学教研室2.中南大学制药工程专业07届毕业生[摘要]本文汇集整理了世界各地寄生于人体的寄生虫608种(不含节肢动物,后者将另行整理),包括原虫116种(含原生动物103种,色虫1种,微孢子虫12种),黏体虫3种,水螅1种,线虫176种,铁线虫23种,环节动物10种(水蛭6种、蚯蚓4种),蛞蝓5种,棘头虫12种,吸虫183种,绦虫76种,涡虫3种。
文中尽量采用最新、有效的生物分类系统给出虫种的分类阶元,并且尽可能列出中文译名,以其为基础和临床寄生虫学工作者提供有价值的参考。
[关键词] 人体,寄生虫,虫种,分类学Species and Biological Taxonomy of Human Parasites in the WorldZHANG Jin-Shun , Zhang Ying-Qian(Department of Parasitology, Hebei North University, Zhangjiakou 075029, China)[Abstract]Understanding systematically the parasites in human is a very important basis for prophylaxis and therapy of parasitc diseases. This article collectes the parasites discovered in persons in 608 species (arthropods were not included)including 116 protozoan(including 103 in Kindom Protozoa, 1 in Kindom Chromista, 12 in Phylum Microspora), 3 myxosoma, 1 hydra, 176 nematodes, 23 horsehair worms,10 annelida(4 earthworms, 6 leeches),5 slugs,12 thorny -headed worms, 183 trematodes, 76 cestodes, 3 turbellarian. To the greatest extent, the valid ranks of taxonomy were used. The Chinese translated terms were gaven as possible as we can for convenience use for basic and clinical researchers.[Key Words] Human,Parasite,Species,Taxonomy人体感染的寄生虫包括寄生的原生动物和蠕虫(线虫、绦虫、吸虫和棘头虫),也包括一些节肢动物、环节动物和软体动物[1]。
虽然有文献推测寄生人体的蠕虫有340多种[2],原虫有80多种[1],但到底从人体发现的寄生虫有多少种?到目前为止,尚缺乏较全面的介绍全球范围内人体寄生虫虫种并反映寄生虫最新生物学分类的资料。
我国已见于文献的人体寄生虫,1997年以前有230种之多[3],以后又陆续有所发现。
瞿逢伊等对我国1871-2006年间首例报告的寄生虫进行了系统的回顾与述评[4]。
然而,国内人体寄生虫学研究存在两大薄弱环节:一是主要研究国内流行的寄生虫,对国际上其他地域流行的寄生虫研究很少,许多见于文献的虫种没有国内资料;二是对寄生虫的生物学分类研究较为落后,仍沿用26年前的分类系统,有些国际上已经废弃的分类阶元仍在使用,而一些已被广泛接受的新的分类词汇还不为大家所熟知[5,6,7]。
此外,国内发现的某些虫种由于文字问题,国外尚未了解。
因此,国内寄生虫病工作者经常会遇到一个问题是,有大量寄生虫学名词不知如何翻译,有许多虫种不知道属于什么分类地位。
WHO“2007年世界卫生报告”指出“现今快速流动的、相互依赖和相互关联的世界为传染病的快速传播、核放射以及有毒物质的威胁创造了无数机会”,“传染病从地理学角度上讲比历史任何时候传播的速度都要快。
据估计,2006年有21亿人次的航空旅客;所以世界上任何一个地方一旦发生疾病暴发或流行,那么仅仅几小时后就会使其他地区大难临头。
”[8]我国随着对外开放和全球一体化进程的加快,国内外交流人员迅猛增加,“根据我国有关部门的统计,2005年中国内地居民出境人数达3103万人次,外国人入境人数为2026万人次,”[9]并有逐年增高的趋势。
与此同时,动、植物及有关物流也十分巨大,这些都为寄生虫病的转播提供了条件。
事实上,近年我国不断发现出国人员将国外流行的寄生虫病带到国内[10,11],因此仅了解我国流行的寄生虫,而不了解世界各地人体感染的寄生虫及其分类学进展对于临床寄生虫病的诊断和鉴别诊断十分不利,与我国社会的发展也是不相适应的。
笔者曾概述了生物分类学的进展,并介绍了Cox(2003)的人体寄生虫分类系统[4]。
然而,Cox的分类仅列出虫种194种,有多种原虫、吸虫、线虫、绦虫未列出,并且也未涉及近年新发现并得到证实的某些虫种、以及可寄生人体的环节动物和软体动物,如鼻孢子虫、新孢子虫、铁线虫、蛭、蛞蝓、蚯蚓、涡虫等[12,13,14,15]。
此外,Cox在分类中原虫省略了科,线虫省略了目,也给参考带来不便。
本文用数年时间,广泛收集国内外文献,特别是有关人体寄生虫病的首发病例的文献[4,16,17,18]、寄生虫综述性文献[19,20,21,22,23]、国内外人体感染性疾病病原体汇总的资料[24,25,26,27]、并参考国内外近年出版的重要专著[28,29,30,31,32,33,34],浏览了众多寄生虫专业网站,对人体发现的608种寄生虫(不含节肢动物,后者将另行整理)进行了汇集整理,其中包括原虫116种(原生动物103种,色虫1种,微孢子虫12种),黏体虫3种,水螅1种,线虫176种,铁线虫23种,环节动物10种(水蛭6种、蚯蚓4种),蛞蝓5种,棘头虫12种,吸虫183种,绦虫76种,涡虫3种。
对于同物异名的虫种,仅按一种计入。
文中尽量采用最新、有效的生物学分类,给出界、门、纲、目、科和种的分类阶元(表1)。
除少数目前因缺乏资料确实难以翻译而仅列出拉丁学名外,尽量将所列虫种给出中文译名,以便寄生虫学工作者参考。
表1 人体感染寄生虫及其生物学分类——————————————————————————————————————————原生动物界(KINGDOM PROTOZOA)原始动物亚界1(Subkingdom I Archezoa)后滴门(Phylum Metamonada)(肠鞭毛虫intestinal flagellates)双滴纲(Class Trepomonadea)双滴目(Order Diplomonadida)六鞭毛科(Family Hexamitidae)蓝氏贾第鞭毛虫(Giardia lamblia同物异名:肠贾第鞭毛虫)肠滴目(Order Enteromonadida)肠滴虫科(Family Enteromonadidae)人肠滴虫(Enteromonas hominis)曲滴纲(Class Retortamonadea)曲滴目(Order Retortamonadida)曲滴虫科(Family Retortamonadidae)迈氏唇鞭毛虫(Chilomastix mesnili),肠旋滴虫(Retortamonas intestinalis),中华内滴虫(R. sinensis)副基体门(Phylum Parabasalia)毛滴纲(Class Trichomonadea)(肠和与肠相关的鞭毛虫intestinal and related flagellates)毛滴目(Order Trichomonadida)毛滴虫科(Family Trichomonadidae)脆弱双核阿米巴(Dientamoeba fragilis),人五毛滴虫(Pentatrichomonas hominis),人毛滴虫(Trichomonas hominis),口腔毛滴虫(T.tenax),阴道毛滴虫(T. vaginalis)动鞭毛纲(Class Zoomastigophorea 或:超鞭毛纲Class Hypermastigida)超鞭毛目(Order Hypermastigida)缨滴虫科(Family Lophomonadae)蠊缨滴虫(Lophomomas blattarum)新生动物亚界2(Subkingdom 2 Neozoa)盘脊下界1(Infrakingdom 1 Discicristata)透色动物门(Phylum Percolozoa)异叶足纲(Class Heterolobosea)(有鞭毛的阿米巴flagellated amoebae)裂核目(Order Schizopyrenida)瓦氏科(Family Vahlkamphidae)福氏耐格里阿米巴(Naegleria fowleri),N. grubery**,Vahlkamplia punctata**,Vahlkamplia lobospinosa**眼虫门(Phylum Euglenozoa)动基体纲(Class Kinetoplastea)(有动基体的鞭毛虫kinetoplastid flagellates)锥体目(Order Trypanosomatida)锥体科(Family Trypanosomatidae)埃塞俄比亚利什曼原虫(Leishmania aethiopica),亚马逊利什曼原虫(L. amazonensis),巴西利什曼原虫(L. braziliensis),哥伦比亚利什曼原虫(L. colombiensis),杜氏利什曼原虫(L. donovani),盖氏利什曼原虫(L. garnhami),圭亚那利什曼原虫(L. guyanensis),婴儿利什曼原虫(L.infantum同物异名:恰氏利什曼原虫L.chagasi),蓝氏利什曼原虫(L. lainsoni),硕大利什曼原虫(L. major),墨西哥利什曼原虫(L. mexicana),奈氏利什曼原虫(L. naiffi),巴拿马利什曼原虫(L. panamensis),秘鲁利什曼原虫(L.peruviana),皮氏利什曼原虫(L. pifanoi),肖氏利什曼原虫(L. shawi),热带利什曼原虫(L. tropica),委内瑞拉利什曼原虫(L. venezuelensis),克氏锥虫(Trypanosoma cruzi),布氏锥虫冈比亚亚种(T. brucei gambiense),布氏锥虫罗德西亚亚种(T. brucei rhodesiense),让氏锥虫(T. rangeli)肉鞭毛下界2(Infrakingdom 2 Sarcomastigota)阿米巴门(Phylun Amoebozoa)叶足亚门(Subphylum Lobosa)阿米巴纲(Class Amoebaea)棘足目(Order Acanthopodida)棘阿米巴科(Family Acanthamoebidae)阿斯特罗尼棘阿米巴(Acanthamoeba astronyxis),堡棘棘阿米巴(A. castellani),卡氏棘阿米巴(A. culbertsoni),A.divionensis,A. griffini,哈氏棘阿米巴(A. hatchetti),A. healyi,A. lugdunensis ,A. mauritaniensis,A. palestinensis,多噬棘阿米巴(A. polyphaga),A. quina,皱棘阿米巴(A. rhysodes),狒狒巴拉姆希阿米巴(Balamuthia mandrillaris),透明哈氏阿米巴(Hartmanella hyaline)**,双核匀变虫(Sappinia diploidea)锥足亚门(Subphylum Conosa)内阿米巴纲(Class Entamoebidea)内阿米巴目(Order Entamoebida)内阿米巴科(Family Entamoebidae)夏氏内阿米巴(Entamoeba chattoni同物异名:波列基内阿米巴E. polecki),结肠内阿米巴(E. coli),迪斯帕内阿米巴(E. dispar),齿龈内阿米巴(E. gingivalis),哈氏内阿米巴(E. hartmanni),溶组织内阿米巴(E.histolytica),茂氏内阿米巴(E. moshkovskii),微小内蜒阿米巴(Endolimax nana),布氏嗜碘阿米巴(Iodamoeba buetschlii)领动门(Phylum Choanozoa)中生粘菌虫纲(Class Mesomycetozoea)肤孢目(Order Dermocystida)鼻孢子虫科(Family Rhinosporideacae )西伯鼻孢子虫(Rhinosporidium seeberi)囊泡下界3(Infrakingdom 3 Alveolata)孢子虫门(Phylum Sporozoa)(孢子虫sporozoans)球虫纲(Class Coccidea)艾美目(Order Eimeriida)艾美科(Family Eimeriidae)人隐孢子虫(Cryptosporidium hominis),安氏隐孢子虫(C.andersoni),猫隐孢子虫(C. felis),火鸡隐孢子虫(C. meleagridis),犬隐孢子虫(C. canis),猪隐孢子虫(C.suis),鼠隐孢子虫(C. muris),微小隐孢子虫(C.parvum),卡耶塔圆孢子虫(Cyclospora cayetanensis),人艾美尔球虫(Eimeriahomonis),贝氏等孢球虫(Isospora belli),纳塔尔等孢球虫(I. natalensis),犬新孢子虫(Neospora caninum),人肉孢子虫(Sarcocystis hominis),林氏肉孢子虫(可能含多个种)(S. lindermanni),猪人肉孢子虫(S.suihominis),刚地弓形虫(Toxoplasma gondii)梨形目(Order Piroplasmida)巴贝科(Family Babesiidae)牛双芽巴贝虫(Babesia bigemina),牛巴贝虫(B. bovis),犬巴贝虫(B. canis),马巴贝虫(B. equi),分歧巴贝虫(B. divergens),吉氏巴贝虫(B. gibsoni),微小(田鼠)巴贝虫(B. microti)泰勒科(Family Theileriidae)小泰勒梨浆虫(Theileria parva)血孢目(Order Haemosporida)疟原虫科(Family Plasmodiidae)巴西疟原虫(Plasmodium brasilianum),食蟹猴疟原虫(P. cynomolgi),食蟹猴疟原虫巴氏种(P. c. bastianelli),艾氏疟原虫(P. eylesi),恶性疟原虫(P. falciparum),猪尾猴疟原虫(P. inui),猪尾猴疟原虫肖氏种(P. i.shortti),诺氏疟原虫(P. knowlesi),三日疟原虫(P. malariae),卵形疟原虫(P. ovale),许氏疟原虫(P. schwetzi),吼猴疟原虫(P. simium),间日疟原虫(P. vivax)纤毛虫门(Phylum Ciliophora)(纤毛虫Ciliates)直口纲(Class Litostomatea)胞口目(Order Vestibulifera)肠袋科(Family Balantidiidae)结肠小袋纤毛虫(Balantidium coli)层咽纲(Class Phyllopharyngea)管口目(Cyrtophorida 或:Order Chlamydodontida)斜管科(Family Chilodonellidae)斜管纤毛虫(Chilodonella spp.)色虫界(KINGDOM CHROMISTA)色物亚界(Subkingdom Chromobiota)双环门(Phylum Bigyra)芽囊纲(Class Blastocystea)芽囊目(Order Blastocystida)芽囊科(Family Blastocystidae)人芽囊原虫(Blastocystis hominis)地位未确定(INCERTAE CEDIS 或:真菌界KINGDOM FUNGI)微孢门(Phylum Microspora)(微孢子虫Microsporidians)微孢纲(Class Microsporea)微孢目(Order Microsporida)脑微孢科(Family Encephalitozoonidae)兔脑微孢虫(Encephalitozoon cuniculi),海伦脑微孢虫(E. hellem),肠脑微孢虫(E. intestinalis同物异名:肠有膈微孢虫Septata intestinalis)肠微孢科(Family Enterocytozoonidae)比氏肠微孢虫(Enterocytozoon bieneusi)微粒科(Family Nosematidae)康氏短粒虫(Brachiola connori原称:人小孢子虫Nosema connori),小泡短粒虫(B. vesicularum),眼微粒虫(Nosema ocularum),角膜条微孢虫(Vittaforma corneae)普孢科(Family Pleistophoridae)害人气管普孢虫(Trachipleistophora anthropophthera),人气管普孢虫(T. hominis)未定非洲微孢虫(Microsporidium africanum),锡兰微孢虫(M. ceylonensis)动物界(KINGDOM ANIMALIA)辐射对称动物亚界1(Subkingdom 1 Radiata)黏体亚界2(Subkingdom 2 Myxozoa)粘体门(Phylum Xyxozoa)粘孢子纲(Class Myxosporea)双壳目(Order Bivalvulida)粘体科(Family Myxozomatidae)鲑鱼尾孢子虫(Henneguya salminicala),尾孢子虫(Henneguya spp.),鸵背鳜碘泡虫(Myxobolus plectrolites)腔肠动物下界(Infrakingdom Coelenterata)刺胞动物门(Phylum Cnidaria)水母亚门(Subphylum Medusozoa)水螅纲(Class Hydrozoa)水螅纲1种(Hydrozoa spp. )侧称亚界3(Subkingdom 3 Bilateria)蜕皮下界1(Infrakingdom 1 Ecdysozoa)线形动物门(Phylum Nemathelminthes)(线虫下门Infraphylum Nematoda,线虫roundworms)有腺纲(Class Adenophorea)(无尾感器纲Aphasmidea)嘴刺目(Order Enoplida)旋毛总科(Superfamily Trichinelloidea)旋毛科(Family Trichinellidae)布氏旋毛虫(Trichinella britovi),米氏旋毛虫(T.murrelli),乡土旋毛虫(T. nativa),纳氏旋毛虫(T. nelsoni),巴布亚旋毛虫(T.papuae),伪旋毛虫(T. pseudospiralis),旋毛形线虫(T. spiralis),T. T5,津巴布韦旋毛虫(T. zimbabwensis)鞭形科(Family Trichuridae)猿猴皮肤线虫(Anatrichosoma cutaneum),肝毛细线虫(Capillaria hepaticum同物异名:Calodium hepaticum,Hepaticola hepatica),菲律宾毛细线虫(C. philippinensis同物异名:Aonchotheca philippinensis),嗜气优鞘线虫(Eucoleus aerophilius同物异名:嗜气毛细线虫Capillaria aerophilius),Pearsonema plica(同物异名:Capillaria plica),鼠鞭虫(Trichuris muris)*,猪鞭虫(T. suis),毛首鞭形线虫(T. trichiura),狐鞭形线虫(T. vulpis)膨结科(Family Dioctophymidae)胃瘤线虫(真圆线虫)(Eustrongylides spp.),肾膨结线虫(Dioctophyma renale)姆斯帕目(Order Muspiceida)姆斯帕总科(Superfamily Muspiceoidea)罗氏科(family Robertdollfusidae)Haycocknema perplexum索虫目(Order Mermithida)索虫科(Family Mermithidae)黑色索线虫(Mermis nigrescens),Agamomermis hominisoris+,Agamomermis restiformis+分肠纲(Class Secernentea)(尾感器纲Phasmidea)蛔目(Order Ascaridida)蛔总科(Superfamily Ascaridoidea)蛔科(Family Ascarididae)似蚓蛔线虫(Ascaris lumbrioides),猪蛔虫(A. suum),浣熊贝蛔虫(Baylisascaris procyonis),网纹前盲囊线虫(Porrocaecum reticulatum),小兔唇蛔虫(Lagochilascaris minor),马副蛔虫(Parascaris equorum),犬弓首线虫(Toxocara canis),猫弓首线虫(T. cati 同物异名:T. mystax),小牛弓首线虫(T.vitulorum)*,山猫弓首蛔虫(T. lyncus)*,马来西亚弓首线虫(T. malaysiensis)*,狐蝠弓首线虫(T. pteropodis)*,狮弓蛔虫(Toxoscaris leonina),钩状宫脂虫(Hysterothylacium aduncum),鲔蛔线虫(Thynnascaris spp.)*异尖科(Family Anisakidae)海异尖线虫(Anisakis marina),简单异尖线虫(A.simplex),抹香鲸异尖线虫(A. physeteris),典型异尖线虫(A. typica),小吻反盲端(接合对盲囊)线虫(Contracaecum osculatum),迷惑伪新地线虫(Pseudoterranovadecipiens同物异名:Phocanema decipiens,Terranova decipiens)圆线目(Order Strongylida)钩口总科(Superfamily Ancylostomatoidea)钩口科(Family Ancylostomatidae)巴西钩口线虫(Ancylostoma braziliense),犬钩口线虫(A. caninum),锡兰钩口线虫(A. Ceylanicum),十二指肠钩口线虫(A.duodenale),日本钩口线虫(A. japonica),马来亚钩虫(A. malayanum),管形钩虫(A. tubaeforme),牛仰钩虫(Bunostomum phlebotomum),羊仰钩虫(B.trigonocephalum),美洲板口线虫(Necator americanus),阿根廷板口线虫(N.argentinus)*,猪板口线虫(N. suillus),狭头弯口线虫(Uncinaria stenocephala)后圆总科(Superfamily Metastrongyloidea)后圆科(Family Metastrongylidae)长后圆线虫(Metastrongylus elongates同物异名: 野猪后圆线虫M. apri,Choerostrongylus),短阴后圆线虫(M. pudendotectu),萨氏后圆线虫(M. salmi)管圆科(Family Angiostrongylidae)广州副圆线虫(Parastrongylus cantonensis 同物异名:广州管圆线虫Angiostrongylus cantonensis),哥斯达黎加副圆线虫(P.Costaricensis 同物异名:哥斯达黎加管圆线虫A. Costaricensis),马克拉斯管圆线虫(A. mackerrasae)*,马来管圆线虫(A. malaysiensis)*毛圆总科(Superfamily Trichostrongyloidea)毛圆科(Family Trichostrongylidae)捻转血矛线虫(Haemonchus contortus),马氏马歇尔线虫(Marshallagia marshalli),指形长刺线虫(Mecistocirrus digitatus),畸形细颈线虫(Nematodirus abnormalis),环纹奥斯特线虫(Ostertagia circumcincta 同物异名:环束背带线虫Teladorsagia circumcincta),奥氏奥斯特线虫(O. ostertagi),近似毛圆线虫(Trichostrongylus affinis),艾氏毛圆线虫(T. axei同物异名:T.extenuatus),短毛圆线虫(T. brevis),有刺毛圆线虫(T. calcaratus),山羊毛圆线虫(T. capricola),蛇形毛圆线虫(T. colubriformis,同物异名:T. instabilis),勒鲁毛圆线虫(T. lerouxi),东方毛圆线虫(T. orientalis),突尾(枪形)毛圆线虫(T.probolurus),斯氏毛圆线虫(T. skrjabini),透明(脆弱)毛圆线虫(T. vitrinus)圆线总科(Superfamily Strongyloidea)夏伯特科(Family Chabertiidae)鼠环齿口钩虫(Cyclodontostomum purvisi同物异名:Ancistronema coronatum),猴结节线虫(帽口食道口线虫)(Oesophagostomum apiostomum同物异名:有刺食道口线虫O. aculeatum),二叉结节线虫(二叉食道口线虫)(O. bifurcum),冠口(猩猩)食道口线虫(O. stephanostomum),缩小三齿线虫(Ternidens diminutus),小三齿线虫(Triodontophorus minor)比翼科(Family Syngamidae)港归比翼线虫(Mammomonogamus gargguiensis),大比翼线虫(M. ingens),喉兽比翼线虫(M. laryngeus),鼻比翼线虫(M. nasicola)尖尾目(Order Oxyurida)尖尾总科(Superfamily Oxyuroidea)尖尾科(Family Oxyuridae)格氏蛲虫(Enterobius gregorii),蠕形住肠线虫(E.vermicularis),隐藏管状线虫(鼠蛲虫)(Syphacia obvelata)杆形目(Order Rhabditida)小杆总科(Superfamily Rhabditoidea)小杆科(Family Rhabditidae)Cheilobus quadrilabiatus(同物异名:Rhabditis schneideri)+,冠双杆线虫(Diploscapter coronata)+,破坏微线虫(Micronema detetrix 同物异名:Halicephalobus gingivalis),土壤拟圆虫(Pelodera strongyloides+同物异名:类圆小杆线虫Rhabditis strongyloides),土壤横形虫(P. transfuga),艾氏小杆线虫(Rhabditis axei)*,R. donbass,长小杆线虫(R. elongata),R. gracilis,人小杆线虫(R. hominis),R. inermis,R. niellyi*,鼠小杆线虫(R. pellio)+,夏赫小杆线虫(R. schachtiella),夏克特小杆线虫(R. schactiella),P. teres+,R. terricola+,R.taurica+,类圆科(Family Strongyloididae)福氏类圆线虫(Strongyloides fuelleborni),S. grasse,河狸鼠类圆线虫(S. myopotami),乳突类圆线虫(S. papillosus),浣熊类圆线虫(S.procyonis),兰氏类圆线虫(S. ransomi),粪类圆线虫(S. stercoralis),韦氏类圆线虫(S. westeri)头叶亚目(Suborder Cephalobina)全凹总科(Superfamily Panagrolaimoidea)全凹科(Family Panagrolaimidae)醋线虫(Turbatrix aceti)+旋尾目(Order Spirurida)龙线总科(Superfamily Dracunculoidea)龙线科(Family Dracunculidae)徽记龙线虫(Dracunculus insignis),麦地那龙线虫(D.medinensis)柔线科(Family Habronematidae)家蝇柔线(马胃)虫(Habronema musca),小口柔线(马胃)虫(Habronema microstoma),大口柔线(马胃)虫(Habronema megastoma)丝虫总科(Superfamily Filarioidea)盘尾科(Family Onchocercidae)Agamofilaria Georgiana,比氏布鲁丝虫(Brugia beaveri)*,圭亚那布鲁丝虫(B. guyanensis),马来布鲁线虫(B. malayi),膨亨布鲁丝虫(B.pahangi),帝汶布鲁线虫(B. timori),樱红唇棘线虫(Dipetalonema arbuta),隐形棘唇线虫(D. reconditum),链尾唇棘线虫(D. streptocerca),Dirofilaria conjunctives,犬恶丝虫(D. immitis),D. magalhesi,匍行恶丝虫(D. repens),条纹恶丝虫(D.striata),皮下丝虫(D. subdermata),细薄恶丝虫(D. tenuis),熊恶丝虫(D. ursi),罗阿丝虫(Loa loa),台湾狐猿线虫(Macacanema formosana),欧氏曼森线虫(Mansonella ozzardi),常现曼森线虫(M. perstans同物异名:常现唇棘线虫Dipetalonema perstans),罗氏曼森线虫(M. rodhaini),半明曼森线虫(M.semiclarum),链尾曼森线虫(M. streptocerca),猴脑膜线虫(Meningonema peruzzii),波利维亚微丝蚴(Microfilaria bolivarensis),M. powelli,旋盘尾线虫(Onchocerca volvulus),班氏吴策线虫(Wuchereria bancrofti),路氏吴策线虫(W. lewisi)丝状科(Family Setariidae)马丝状线虫(Setaria equina)颚口总科(Superfamily Gnathostomatoidea)颚口科(Family Gnathostomatidae)双核鄂口线虫(Gnathostoma binucleatum),多氏鄂口线虫(G. doloresi),刚刺颚口线虫(G. hispidum),马来西亚鄂口线虫(G.malaysiae),日本鄂口线虫(G. nipponicum),棘颚口线虫(G. spinigerum),G. vivarina 旋尾总科(Superfamily Spiruroidea)筒线科(Family Gongylonematidae)美丽筒线虫(Gonglonema pulchrum)尾旋科(Family Spirocerciden)狼尾旋线虫(血色食道虫)(Spirocerca lupi 同物异名:Spirocerca sanguinolenta)华首科(锐形科)(Family Acuariidae)扭头唇饰带线虫(Cheilospirura hamulosa)四棱科(Family Tetrameridae)裂刺四棱线虫(Tetrameres fissispinus)吸吮总科(Superfamily Thelazioidea)吸吮科(Family Thelaziidae)加利福尼亚吸吮线虫(Thelazia californiensis),结膜吸吮线虫(T. callipaeda),罗德西吸吮线虫(T. rhodesii)奇口科(Family Rictulariidae)侧翼棘奇口线虫(Rictularia pterygodermatites)泡翼总科(Superfamily Physalopteroidea)泡翼科(Family Physalopteridae)高加索泡翼线虫(Physaloptera caucasica),横形泡翼线虫(P. transfuga)垫刃目Tylenchida垫刃总科(Superfamily Tylenchoidea)垫刃科(Family Tylenchidae)鳞球茎茎线虫(Ditylenchus dispaci),根瘤线虫(Ditylenchus radicicola同物异名:Meloidogyn marioni M. javanica,Heterodera radicicola,H.marioni)线形虫动物门(Phylum Nematomorpha)铁线虫纲(Class Gordioida)铁线虫目(Order Gordioidea)铁线虫科(Familly Gordiidae)水生铁线虫(Gordius aquaticus),智利铁线虫(G. chilensis),G. gesneri,G. inesae,G. ogatai,G. perroneiti,G. skorikowi,粗壮铁线虫(G.robustus同物异名:绒毛铁线虫G. villoti),刚节铁线虫(G. setiger)粗皮目(Order Chordodea)粗皮科(Familly Chordodidae)Chordodes capensis*拟粗皮科(Family Parachordodidae)哥伦比亚新粗皮虫(Neochordodes colombianus)*,Parachordodes alpestris*,P. pustulosus*,P. raphaelis,P. tolusanus*,P. violaceus*,P. wolterstoffi拟铁线科(Family Paragordiidae)Paragordius areolatus*,P. cinctus*,P. esavianus,P. tricuspidatus*,P. varius*,Pseudogordius tanganyikae*前口类(Branch Protostomia)环节动物门(Phylum Annelida)蛭纲(Class Hirudinea)(蚂蟥Leeches)真蛭亚纲(Subclass Euhirudinea)无吻蛭目(Order Arhynchobdellida)医蛭形亚目(Suborder Hirudiniformes)医蛭科(Family Hirudinidae)凶恶怖蛭(鼻蛭)(Dinobdella ferox),尼罗沼蛭(Limnatis nilotica)山蛭科(Family Haemadipsidae)日本医蛭(Haemadipsa japonica),蒙大拿山蛭(H.montana),森林山蛭(H. sylvestris),锡兰山蛭(H. zeylanica)寡毛纲(Class Oligochaeta)单向蚓目(Order Haplotaxida)(蚯蚓Earthworm)巨蚓科(Family Megascolecidae)异毛环毛蚓(Pheretima diffringens)正蚓科(Family Lumbricidae)赤子爱胜蚓(Eisenia foetida)颤蚓目(Order Tubificida )线蚓科(Family Enchytraeidae)白线蚓(Fridericia spp.)仙女虫科(Family Naididae)仙女虫(Nais spp.)软体动物门(Phylum Mollusca)腹足纲(Class Gastropoda)柄眼目(Order Stylommatophora)蛞蝓科(Family Limacidae)瓦仑西亚那蛞蝓(Lehmannia valentiana),蛞蝓(Limax spp.)蠕蝓科(Family Boettgerillidae)野蛞蝓(光滑蛞蝓)(Agriolimax agrestis)肺螺亚纲(Subclass Pulmonata)鞋形目(Order Soleolifera)复套蛞蝓科(Family Veronicellidae,或:足壁蛞蝓科Vaginulidae)双线黏液蛞蝓(Philomycus bilinetus),玉溪复套蛞蝓(Vaginulus yuxisis sp. nov)扁形下界2(Infrakingdom 2 Platyzoa)棘颚门(Phylum Acanthognatha)(头部多刺的蠕虫thorny-or spiny-headed worms)原棘纲(Class Archiacanthocephala)念珠目(Order Moniliformida)念珠科(Family Moniliformidae)念珠棘头虫(Moniliformis moniliformis),M. dubius,寡棘吻目(Order Oligacanthorhynchida)寡棘吻科(Family Oligacanthorhynchidae)猪巨吻棘头虫(Macracanthorhynchus hirudinaceus),大巨吻棘头虫(M. ingens)古棘纲(Class Palaeacanthocephala)棘吻目(Order Echinorhynchida)棘吻科(Family Echinorhynchidae)蟾酴棘头虫(Acanthocephalus bufonis),A. marcianae,饶氏棘头虫(A. rauschi),隐形棘头虫(A. recondita),蟾酴伪棘头虫(Pseudoacanthocephalus bufonis)+多形目(Order Polymorphida)多形科(Family Polymorphidae)球茎体虫(钩头虫类)(Bolbosoma spp.),Corynosoma strumosum+始新棘头虫纲(Class Eoacanthocephala)圆棘头目(Order Gyracanthocephala)四旋棘科(Family Quadrigyridae)隐棘新棘体虫六安亚种(Neosentis celatus liuanensis)扁形动物门(Phylum Platyhelminthes)复殖纲(Class Digenea)(吸虫纲Trematoda,吸虫flukes)鸮形目(Order Strigeida)双穴科(Family Diplostomidae)美洲重翼吸虫(Alaria americana),皱缩重翼吸虫(A.marcianae),篦状(匙形)复口吸虫(Diplostomum spathaceum),新双穴科(Family Neodiplostomidae)杯状海狸吸虫(Fibricola cratera)#,汉城新复口吸虫(Neodiplostomum seoulense同物异名:汉城海狸吸虫Fibricola.seoulensis)裂体科(Family Schistosomatidae)特瑞哥澳毕吸虫(Austrobilharzia terrigalensis),波兰小毕吸虫(Bilharziella Polonica),Giganobilharzia huttoni,椋鸟巨毕吸虫(G.sturniae),美洲异毕吸虫(Heterobilharzia americana),微毕吸虫(Microbilharziaspp.),程氏东毕吸虫(Orientobilharzia cheni),土耳其斯坦东毕吸虫(O.turkestanica),土耳其斯坦东毕吸虫结节亚种(O. t. tuberculata),土耳其斯坦鸟毕吸虫(Ornithobilharzia turkestanicum),牛裂体吸虫(Schistosoma bovis),卡图裂体吸虫(S. cattoi),埃及血吸虫(S. haematobium),S. incognitum*,间插血吸虫(S. intercalatum),日本血吸虫(S. japonicum),马来血吸虫(S. malayensis),曼氏血吸虫(S. mansoni),马氏血吸虫(S. margrebowei),梅氏裂体吸虫(S.mattheei),湄公血吸虫(S. mekongi),菲律宾裂体吸虫(S. philippinensis)*,罗氏裂体吸虫(S. rodhaini),中华血吸虫(S. sinensium),梭形裂体吸虫(S.spindale)*,S. sindale,杜氏小裂体吸虫(Schistosomatium douthitti),短毛毕吸虫(Trichobilharzia brevis),大榆树毛毕吸虫(T. dayushuensis),巨大毛毕吸虫(T. gigantea),广东毛毕吸虫(T. guangdongenis),集安毛毕吸虫(T. jianensis),眼点毛毕吸虫(T. ocellata),包氏毛毕吸虫(T. paoi),平南毛毕吸虫(T. pingnana),沼栖毛毕吸虫(T. stagnicolae),中山毛毕吸虫(T. zhongshani)弯口科(Family Clinostomatidae)平坦弯口吸虫(Clinostomum complanaturn)等睾科(Family Isoparorchidae)鳔等睾吸虫(Isoparorchis hypselobagri)裸茎科(Family Gymnophallidae)徐氏拟裸茎吸虫(Gymnophalloides seoi)杯叶科(Family Cyathocotylidae)活动前半口吸虫(Prohemistomum vivax)鸮形科(Family Strigeide)日本杯尾吸虫(Cotylurus japonicus)短咽科(Family Brachylaimidae)断壳蜗短咽吸虫(Brachylaima ruminate)棘口目(Order Echinostomatida)棘口科(Family Echinostomatidae)马来亚刺口吸虫(Artyfechinostomum malayanum),梅氏刺口吸虫(A. mehrai),狭睾棘隙吸虫(Echinochasmus augustitestis),福建棘隙吸虫(E. fujianensis),日本棘隙吸虫(E. japonicum),九佛棘隙吸虫(E.jinfoensis),藐小棘隙吸虫(E. liliputanus同物异名?:藐小棘口吸虫Echinostomaliliputanus),Echinoparapyphium paraulum*,埃及棘口吸虫(Echinostomaaegyptica),移睾棘口吸虫(E. cinetorchis同物异名?:移睾棘隙吸虫Echinochasmus cinetorchis),多刺(外旋)棘口吸虫(E. echinatum同物异名:林杜棘口吸虫E. lindoensis),圆圃棘口吸虫(E. hortense),日本棘口吸虫(E.japonicus),巨睾棘口吸虫(E. macrorchis),马来亚棘口吸虫(E. malayanum同物异名:马来亚真缘吸虫Euparyphium malayanum),宫川棘口吸虫(E.miyagawai),接睾棘口吸虫(E. paraulum),叶形棘口吸虫(E. perfoliatus 同物异名?:抱茎棘隙吸虫或叶形棘隙吸虫Echinochasmus perfoliatus),反弯棘口吸虫(E. recurvatum同物异名:曲颈棘缘吸虫Echinoparyphium recurvatum,Echinoparyphium koidzumii),卷棘口吸虫(E. revolutum),犬外隙吸虫(Episthmiumcaninum),伊族真缘吸虫(Euparyphium ilocanum 同物异名:伊族棘口吸虫Echinostoma ilocanum),雅西真缘吸虫(E. jassyense同物异名:雅西棘口吸虫Echinostoma jassyense,獾真缘吸虫E. melis),穆氏刺茎(鞭带)吸虫(Himasthlamuehlensi),似锥低颈吸虫(Hypoderaeum conoideum)蹲茎科(Family Cathaemasiidae)叶蹲茎吸虫(Cathaemasia cabrerai)片形科(Family Fasciolidae)巨片吸虫(Fasciola gigantica),肝片形吸虫(F. hepatica),印度片形吸虫(F. indica),布氏姜片吸虫(Fasciolopsis buski)合叶科(Family Zygocorylidae,或:同端盘科Paramphistomatidae)长菲策吸虫(Fischoederius elongatus),人似腹盘吸虫(Gastrodiscoides hominis),瓦氏瓦生吸虫(Watsonius watoni),霍克同盘吸虫(Amphistoma hawkesii)嗜眼科(Family Philophthalmidae)涉禽嗜眼吸虫(Philophthalmus gralli),泪嗜眼吸虫(Philophthalmus lachrymosus或lacrymosus)光睾科(Family Psilostomatidae)人光睾吸虫(Psilorchis hominis)*,多棘缘口吸虫(Paryphostomum sufrartyfex同物异名?:多棘刺口吸虫Artyfechinostomum sufrartyfex)斜睾目(Order Plagiorchiida)双腔科(Family Dicrocoeliidae)中华双腔吸虫(Dicrocoelium chinensis)*,支双腔吸虫(D. dendriticum同物异名:矛形双腔吸虫D. lanceolatum又称“柳叶样吸虫”),牛双腔吸虫(D. hospes),胰阔盘吸虫(Eurytrema pancreaticum),异形科(Family Hererophyidae)齿形背茎吸虫(Apophallus donicus),迷人歧阴茎吸虫(A. venustus),凹管吸虫(Ascocotyle spp.同物异名:Phagicola spp.),多刺棘带吸虫(Centrocestus armatus# 同物异名:多刺瓶状吸虫S. armatus),阿氏棘带吸虫(C. asadai),犬棘带吸虫(C. caninus),尖端棘带吸虫(C. cuspidatus),台湾棘带吸虫(C. formosanus同物异名:台湾瓶状吸虫Stamnosoma formosanus),C. kurokawai,长棘带吸虫(C. longus),舌隐叶吸虫(Cryptocotyle lingua),小睾单睾吸虫(Haplorchis microrchis),H. pleurolophocerca,钩棘单睾吸虫(H. pumilio 同物异名:H. taihokui,Monorchotrema taihokui),扇棘单睾吸虫(H. taichui同物异名:扇棘单睾孔虫M. taichui),横川(多棘)单睾吸虫(H. yokogawai同物异名:横川单睾孔虫Monorchotrema yogogawai),H. vanissimus,短盲肠异形吸虫(Heterophyes brevicaeca),连结异形吸虫(H. continus),异异形吸虫(H. dispar),等异形吸虫(H. equalis),异形异形吸虫(H. heterophyes同种异名:H. nocens),桂田异形吸虫(H. katzuradai),连结拟异吸虫(Heterophyopsis continua),桂田后殖吸虫(Metagonimus katzuradai),微小后殖吸虫(M. minutus),高桥后殖吸虫(M. takahashii),横川后殖吸虫(M. yokogawai同物异名:横川异形吸虫H.yokogawai),哥氏原角囊吸虫(Procerovum calderoni同物异名:哥氏单睾吸虫Haplorchis calderoni,哥氏单睾孔吸虫Monorchotrema calderoni),施氏原角囊吸虫(P. sisoni),变异前角囊吸虫(P. varium)#,前肠肾形吸虫(Pygidiopsis summa),双盲端星隙吸虫(Stellantchasmus amplicaecalis同物异名:双盲端两孔吸虫Diorchitrem amplicaecale),镰刀星隙吸虫(S. falcatus同物异名:镰状两睾孔吸虫Diorchitrema falcatus),台湾星隙吸虫(S. formosanus 同物异名:台湾两睾孔吸虫D. formosanim),伪触毛星隙吸虫(S. pseudocirrata同物异名:伪触毛两睾孔吸虫D. pseudocirrata),Stictodora fuscatum,三枝斑皮吸虫(S. tridactyla)后睾科(Family Opisthorchiidae)微小伪猫对体吸虫(Amphimerus pseudodelineus minutus),白次睾吸虫(Metorchis albidus),胆囊次睾吸虫(M. bilis),结合次睾吸虫(M. conjunctus),东方次睾吸虫(M. orientalis),台湾次睾吸虫(M.taiwanensis),猫后睾吸虫(Opisthorchis felineus同物异名:细颈后睾吸虫O.tenuicollis),瓜亚基尔后睾吸虫(O. guayaquilensis同物异名:伪猫对体吸虫Amphimerus pseudofelineus),继母(犬)后睾吸虫(O. noverca 同物异名:继母(犬)对体吸虫A. noverca,猫对体吸虫A. caninus),华后睾吸虫(O. sinensis 同物异名:华支睾吸虫Clonorchis sinensis),麝猫后睾吸虫(O. viverrini),埃塞俄比亚拟端盘吸虫(Pseudamphistomus aethiopicum),截形拟端盘吸虫(P. truncatum)枝腺科(Family Lecithodendriidae)莫氏拟枝腺吸虫(Paralecithodendrium molenkampi),钝叶拟枝腺吸虫(P. obtusum),具腺拟枝腺吸虫(P. glandulosum),彭氏显茎吸虫(Phaneropsolus bonnei),刺茎显茎吸虫(P. spinicirrus),莫氏前肠腺吸虫(Prosthodendrium molenkampi)并殖科(Family Paragonimidae)非洲并殖吸虫(Paragonimus africanus),亚马逊并殖吸虫(P. amazonicas)*,曼谷并殖吸虫(P. bankokensis)*,卡利并殖吸虫(P.caliensis)*,三平正并殖吸虫(P. cenocopiosus)*,结实并殖吸虫(P. compactus)*,刚果并殖吸虫(P. congolensis),异盘并殖吸虫(P. heterotremus同物异名:团山并殖吸虫P. tuanshenensis),印加并殖吸虫(P. inca)*,克氏并殖吸虫(P.kellicotti可能的同物异名:粗壮并殖吸虫P. rudis),巨睾并殖吸虫(P. macrorchis)*,墨西哥并殖吸虫(P. mexicanus同物异名:厄瓜多尔并殖吸虫P. ecuadoriensis,秘鲁并殖吸虫P. peruvienus),宫崎氏并殖吸虫(P. miyazakii),大平并殖吸虫(P.ohirai同物异名:怡乐存并殖吸虫P. iloktsuenensis,佐渡并殖吸虫P. sadoensis),丰宫并殖吸虫(P. proliferus同物异名:勐腊并殖吸虫P. menglaensis*),肺生并殖吸虫(P. pulmonalis),林氏并殖吸虫(P. ringeri),斯氏并殖吸虫(P. skrjabini 同物异名:四川并殖吸虫P. szechuanensis,会同并殖吸虫P. huitongensis,陈氏并殖吸虫P. cheni,泡囊并殖吸虫P. veocularis,异睾并殖吸虫P. hetrorchis),双侧宫并殖吸虫(P. uterobilateralis),卫氏并殖吸虫(P. westermani 同物异名:艾氏并殖吸虫P. edwardsi,猕猴并殖吸虫P. macacae,扁囊并殖吸虫P.asymmetricus),卫氏并殖吸虫菲律宾亚种(P. w. filipinus 同物异名:菲律宾并殖吸虫P. phillipinensis,菲律宾并殖吸虫P. filipinus),卫氏并殖吸虫伊春亚种(P.w. inchunensis),卫氏并殖吸虫日本亚种(P. w. japonicus),卫氏并殖吸虫四川变种(P. w. var szechuanensis),云南并殖吸虫(P. yunnanensis*可能的同物异名:哈氏并殖吸虫P. harinasuti,小睾并殖吸虫P. microrchis)繁睾科(Family Achillurbainiidae)新繁睾吸虫(Achillurbainia nouveli),隐形繁睾吸虫(A. recondita),刚果变吸虫(Poikilorchis congolensis)斜睾科(Family Plagiorchiidae)Plagiorchis hiranasuta(=P. harinasutai),爪哇斜睾吸虫(P. javanensis(=P. javensis),鼠斜睾吸虫(P. muris),菲律宾斜睾吸虫(P.philippinensis)隐孔科(Family Troglotrematidae 或:Family Nanophyetidae)鲑隐孔吸虫(Nanophyetus salmincola 同物异名:Troglotrema salmincola),N. schikhobalowi 微茎科(Family Microphallidae)短肠微茎吸虫(Microphallus brevicaeca同物异名:Carneophallus brevicaeca,Spelotrema brevicaeca),微小微茎吸虫(M. minus)# 绦虫纲(Class Cestoidea)(多节绦虫纲Cestoda,绦虫tapeworms)假叶目(Order Pseudophyllidea)双叶槽科(Family Diphyllobothriidae)雅西布郎绦虫(Braunia jasseyensis),布氏双线绦虫(Digramma brauni),阿拉斯加裂头绦虫(Diphyllobothrium alascense),卡梅伦裂头绦虫(D. cameroni),心形裂头绦虫(D. cordatum),达勒裂头绦虫(D. dalliae),树突裂头绦虫(D. dendriticum可能的同物异名:D. giljacicum,小裂头绦虫D. minus,斯氏裂头绦虫D. skrjabini,D. tungussicum,熊裂头绦虫D.ursi,S. nenzi),秀丽裂头绦虫(D. elegans),D. hians,克氏裂头绦虫(D.klebanovskii),矛形裂头绦虫(D. lanceolatum),阔节裂头绦虫(D. latum),日本海裂头绦虫(D. nihonkaiense),逆戟鲸裂头绦虫(D. orcini),太平洋裂头绦虫(D. pacificum),短小裂头绦虫(D. parvum),D. pretoriensis,D. scoticum,D. srictum,D. stemmacephalum(同物异名:D. yonagoensis),泰氏裂头绦虫(D.theileri同物异名:泰氏迭宫绦虫S. theileri,S. pretoriensis),大复殖孔绦虫(Diplogonoporus brauni),福岗复殖孔绦虫(D. fukuokaensis),犬复殖孔绦虫(D.grandis 同物异名:D. balaenopterae),肠舌形绦虫(Ligula intestinalis),Pyramicocephalus anthrocephalus,实心裂头绦虫(Schistocephalus solidus),曼氏裂头蚴(Sparganum mansoni),增殖裂头蚴(S. proliforum),巴氏迭宫绦虫(Spirometra baxteri),胡氏迭宫绦虫(S. houghtoni 同物异名:胡氏裂头绦虫D.houghtoni,)曼氏迭宫绦虫(S. mansoni 同物异名:曼氏裂头绦虫D.mansoni,可能的同物异名:猬裂头绦虫D. erinaceieuropaei,D.erinoceiuropae,D. erinacei,猬迭宫绦虫S.erinoceiuropae,S. erinacei),拟曼氏迭宫绦虫(S. mansonoides 同物异名:拟曼氏裂头绦虫D.mansonoides)圆叶目(Order Cyclophyllidea)裸头科(Family Anoplocephalidae)短尖伯特绦虫(Bertiella mucronata),司氏伯特绦虫(B. studeri),对称马带绦虫(Mathevotaenia symmetrica 同物异名:Opussumia,Inversia,Morosovella,Paratriotaenia,Markewitschitaenia,Hickmania,Schizorchodes,Priodontia,Linstoparonia,Mangustella,Vasoramia),扩展莫尼茨绦虫(Moniezia。